Ueckermann, Edward A.¹

¹North-West University School of Environmental Sciences and Development (Zoology), South-Africa.

2021 - Volume: 61 Issue: 1 pages: 201-206

Obituary

Professor Pieter Daniel Theron was born in Britstown, Northern Cape, South Africa in 1945. He matriculated at the age of 16 where after he boarded a train to undertake his studies at the then Potchefstroom University for Christian Higher Education (PU for CHE). Upon completion of his BSc in 1965, he was convinced by the Head of Department, Prof. P.A.J. Ryke to rather pursue postgraduate studies than go into teaching as originally intended. He completed his BSc Honours and MSc in 1968. He received his Doctor Scientiae (DSc) under the supervision of Prof. Ryke based on a taxonomic-ecological study of selected edaphic Endeostimata (Trombidiformes: Acari) in 1972. He was employed in the Department of Zoology at PU for CHE since 1966 until his retirement in 2011. During this period, he was Head of Department of both Zoology and the UNISA Life Sciences that was presented by North-West University (and the former PU-CHE). Even after his retirement he remained active in both teaching and research at the NWU as a Post-65 and extraordinary Professor appointment until 2018.

As researcher, Pieter Theron, specialized in the mite suborder Endeostigmata but towards the end of his career was also involved in projects on other mite families, a total 42 papers, resulting from his collaborations with 35 co-workers. The bulk of his 71 publications were published after 2003. He was instrumental in the description of 112 new species, 23 new genera,and two new families and one new subfamily and tribus. The depictions in his publications were masterly composed. This served as a catalyst for the interest of other researchers in the Endeostigmata. He had a long research relationship with Prof. Yves Coineau of the Muséum National d'Histoire Naturelle, Arthropodes, Paris, France. Their area of interest was the rock-dwelling Adamystidae and Saxidromidae from which five papers resulted and additionally a number of specimens were still in the process of being described. Just before his retirement and thereafter he collaborated with Prof. Nestor Fernandez of the National Council of Scientific and Technological Research, La Rioja University, La Rioja, Argentina on several oribatid families which resulted in 32 papers. At the end of his career, he assisted Russian Acarologists during their visit to South Africa in 2016 and 2018 in their search for mites associated with anthills. From this short collaboration, eight articles resulted and a new species named after Pieter Theron, namely Tanytydeus theroni (Paratydeidae). During his career, seven additional mite species was named after him: Anaplonobia theroni and Neopetrobia theroni (Tetranychidae), Stigmaeus pieteri (Stigmaeidae), Typhlodromus (Anthoseius) theroni (Phytoseiidae), Adamystis theroni (Adamystidae), Grandjeanicus theroni (Grandjeanicidae) and Uropoda theroni (Uropodidae). Pieter Theron was also the first chair-person of the African Acarology Association established in 1998 in Pretoria till 2004.

As teacher and mentor, Pieter Theron will be remembered fondly by all his students and specifically those first-year students (which include numerous current and retired academics) whom he taught over his more than 50 years of teaching at the North-West University (the then Potchefstroom University for CHE). During this time, he was also one of the developers and first presenters of the UNISA Zoology practicals presented in Potchefstroom since the mid-1960s. For decades, he was also involved in the week-long Ecology field excursion of the third-year NWU and UNISA Zoology students. Most of his colleagues will attest to the fact that he could always be approached for valuable advice (and acting as a soundboard) on lecturing or logistical issues with regards to teaching undergraduate modules. Numerous graduated students will attest to the fact that they've known Prof. Theron from their years as school learners. He was always keen and approachable to explain to them and their parents what Zoology as university subject entailed and possible career possibilities to be pursued. After his retirement in 2011 he stayed on as an extraordinary Professor and was still actively involved in teaching undergraduate modules in ''Philosophy of Science'' and as a guest lecturer for Invertebrate Zoology. He was also the main contributor to a publication on the history of the Zoology department, documenting the centenary celebrations of the NWU Faculty of Natural Sciences in 2017.

Even though acarology and taxonomy are not a very popular field of research interest, Pieter was always able to attract postgraduate students due to the personal interest he took in them. In some cases, he would even successfully bet on the academic underdog whom ultimately would make a success of their studies. In this way he made a contribution in their professional and personal development. He lived his research out in teaching by always elucidating with examples from his field work; and even at home, allowing some of his Acari pets (ticks) to feed on his pet dogs to the dismay of his wife. Pieter Theron was a precise scientist with a formidable memory. A memory where he treasured memories of a life time, a lifetime of colleagues who will now remember him with fondness.

Colleagues, North-West University, School of Environmental Sciences and Development (Zoology)

Potchefstroom

South Africa

Articles published by P.D. Theron

References

1. Theron P.D., Ryke P.A.J. 1969. The family Nanorchestidae Grandjean (Acari: Prostigmata) with descriptions of new species from South African soils. Journal of the Entomological Society of southern Africa 32: 31-60.

2. Theron P.D., Meyer M.K.P., Ryke P.A.J. 1970. Two new genera of the family Paratydeidae (Acari: Prostigmata) from South African soils. Acarologia 11: 697-710.

3. Theron P.D., Meyer M.K.P., Ryke P.A.J. 1970. The family Alicorhagiidae Grandjean (Acari: Trombidiformes) with descriptions of a new genus and species from South African soils. Acarologia 12: 668-676.

4. Theron P.D. 1974. Hybalicidae, a new family of Endiostigmatic mites (Acari: Trombidiformes). Acarologia 16: 397-412.

5. Theron P.D. 1975. Two new species of the family Nanorhestidae (Acari: Endeostigmata) from pasture soil in South Africa. Wetenskaplike bydraes van die P. U. for C.H.O. Reeks B 63: 1-9.

6. Theron P.D., Ryke P.A.J. 1975. Three new species of the family Sphaerolichidae (Acari: Endeostigmata) from South Africa. Acarologia 17: 220-235.

7. Theron P.D., Ryke P.A.J. 1975. Five new species of the family Lordalychidae (Acari: Endeostigmata) from South Africa. Acarologia 17: 631-651

8. Theron P.D. 1976. New species of the genus Terpnacarus Granjean (Acari: Terpnacaridae) with notes on the biology of one species. Journal of the Entomological Society of southern Africa 39: 131-141.

9. Theron, P.D. 1977. New species of the genus Petralycus Grandjean (Acari: Endeostigmata) from South Africa. Acarologia 19: 38-45.

10. Theron P.D. 1979. The functional morphology of the gnathosoma of some liquid and solid feeders in the Trombidiformes, Cryptostigmata and Astigmata (Acarina): 575-579. In: E. Piffl (ed.) Proceedings of the 4^th International Congress of Acarology. Akademiai Kiado: Budapest.

11. Jordaan L.C., Loots, G.C., Theron P.D. 1987. New species of the genus Antennoseius Berlese, 1916 (Acari: Ascidae) from the Afrotropical Region, with a redescription of A. boskopensis Ryke. Phytophylactica 19: 237-241.

12. Jordaan L.C., Loots G.C., Theron P.D. 1987. Two new species of the genus Cheiroseius Berlese, 1916 (Acari: Ascidae) from the Afrotropical Region. Phytophylactica 19: 231-235.

13. Jordaan L.C., Loots G.C., Theron P.D. 1988. A new genus and new species of Rhodacaridae (Acari) from the Afrotropical Region. Journal of the Entomological Society of southern Africa. 51(2): 275-281.

14. Olivier P.A.S., Theron P.D. 1989. A new species of Speleorchestes (Nanorchestidae: Prostigmata) from a savanna biotope in South Africa. South African Journal of Zoology 24: 356-360. doi:10.1080/02541858.1989.11448177

15. Loots G.C., Theron P.D. 1992. Adhaerenseius floralis, a new ascid genus and species (Acari: Parasitiformes: Ascidae) associated with Poellnitzia rubriflora (L. Bol) Uitewaal in the Western Cape Province of South Africa. Journal of the Entomological Society of southern Africa 55: 39-49.

16. Olivier P.A.S., Theron, P.D. 1997. The genus Benoinyssus Fain (Acari: Eupodidae) from southern Africa, with descriptions of five new species. African Entomology 5: 301-318.

17. Olivier P.A.S., Theron, P.D. 1997. The genus Eupodes Koch, 1835 (Acari: Prostigmata: Eupodidae) from southern African soil and vegetation. Part I. Characterisation of the genus, designation of the type species and descriptions of three new species. Koedoe 40: 57-73. doi:10.4102/koedoe.v40i2.273

18. Olivier P.A.S., Theron, P.D. 1999. Pentapalipidae, a new family of eupodoid mites (Prostigmata: Eupodoidea) from South Africa. Acarologia 40: 385-392.

19. Van der Schyff J., Theron P.D., Ueckermann E.A. 2003. Polythrichinae, a new subfamily of Bdellidae (Acari: Prostigmata) from the Afrotropical Region. African Plant Protection 9: 19-22.

20. Van der Schyff J., Theron P.D., Ueckermann E.A. 2004. Hexabdella, a new genus of Bdellidae (Acari: Prostigmata) from southern Africa, with descriptions of five new species. African Plant Protection 10: 13-25.

21. Van der Schyff J., Theron P.D., Ueckermann E.A. 2005. Descriptions of seven new species of the genus Bdella Latreille (Acari: Prostigmata: Bdellidae) from southern Africa and a redescription of B. neograndjeani Meyer & Ryke. African Entomology 13: 221-238.

22. Coineau Y., Theron P.D., Fernandez N. 2006. Parades et dimorphismes sexuels compares chez deux nouveau genres de Saxidromidae (Acari, Alycina) d'Afrique du Sud. Acarologia 46: 65-87

23. Omukunda E., Theron P.D., Ueckermann E.A. 2007. Six new species of the genus Biscirus Thor (Acari: Bdellidae) from South Africa. Acarina 15: 269-282.

24. Toroitich F.J., Ueckermann E.A., Theron P.D., Knapp M. 2009. The tetranychid mites (Acari: Tetranychidae) of Kenya and a redescription of the species Peltanobia erasmusi Meyer (Acari: Tetranychidae) based on males. Zootaxa 2176: 33-47. doi:10.11646/zootaxa.2176.1.3

25. Ueckermann E.A., Theron P.D., Tiedt L.R. 2010. Revision of the genus Obuloides Baker and Tuttle, 1975 (Acari: Tenuipalpidae), with the description of five new species. International Journal of Acarology 36: 151-167. doi:10.1080/01647950903555442

26. Alberti G., Coineau Y., Fernandez N., Theron P.D. 2010. On the female genital system of Saxidromidae (Prostigmata: Actinotrichida) XIII International Congress of Acarology, Recife, Brazil, 23-27 August.

27. Fernandez N., Cleva R., Theron P.D. 2010. Idiozetes malgache, a new species of Idiozetidae (Acari: Oribatida) from Madagascar. International Journal of Acarology 36(5): 437-451. doi:10.1080/01647954.2010.485132

28. Fernandez N., Theron P.D., Cleva R. 2011. Rogerzetes lacouturieri gen. nov., n. sp., (Acari: Oribatidae: Eremaeozetidae) from Madagascar. International Journal of Acarology 37(suppl. I): 61-70. doi:10.1080/01647954.2010.539980

29. Fernandez,N., Cleva R. & Theron P.D. 2011. Malgachebates peyrierasi gen. n. sp. (Acari: Oribatida: Plasmobatidae) from Madagascar. International Journal of Acarology 37(1): 61-74. doi:10.1080/01647954.2010.495082

30. Toroitich F.J., Ueckermann E.A., Theron P.D., Knapp M., Haas F. 2011. The genus Brevinychus Meyer, (Acari: Tetranychidae) with the description of a new species from Tanzania. International Journal of Acarology 37(suppl. I): 149-155. doi:10.1080/01647954.2010.543700

31. Omukunda E., Theron P.D., Ueckermann E.A. 2012. Spinibdella Thor (Acari: Bdellidae) from southern Africa: descriptions of five new species and the redescription of S. thori (Meyer & Ryke). Zootaxa. 3304: 1-24. doi:10.11646/zootaxa.3304.1.1

32. Wahl J.J., Theron P.D., Maboeta M.S. 2012. Soil mesofauna as bioindicators to assess environmental disturbance at a platinum mine in South Africa. Ecotoxicology and Environmental Safety 86: 250-260. doi:10.1016/j.ecoenv.2012.09.027

33. Alberti G., Coineau Y., Fernandez N., Theron P. 2012. Fine Structure of the Saxidromid Gnathosoma with Remarks on additional of the Proterosoma (Acari, Actinotrichida) 7th Symposium of the EURAAC (European Association of Acarologists) Vienna, Austria 9-13 July.

34. Du Preez G., Theron P.D. & Fourie D. 2013. Terrestrial mesofauna biodiversity in unique karst environments in southern Africa: 386-390. Proceedings of 16^th International congress of Speleology, Brno, Czech Republic, 21-28 July.

35. Fernandez N., Theron P.D., Rollard C. 2013. Revision of the family Carabodidae (Acari: Oribatida) I. Description of a new genus, Bovicarabodes with three new species, and the redescription of Hardybodes mirabilis Balogh, 1970. International Journal of Acarology 39(1): 26-57. doi:10.1080/01647954.2012.741144

36. Fernandez N., Theron P.D., Rollard C., Leiva S. 2013. Revision of the family Carabodidae (Acari: Oribatida) II. Redescription of Austrocarabodes ensifer (Sellnick, 1931), Aokiella florens Balogh & Mahunka, 1967 and Singabodes rarus Mahunka, 1998. International Journal of Acarology 39(3): 181-199. doi:10.1080/01647954.2012.754493

37. Fernandez N., Theron P.D., Rollard C., Leiva S. 2013. Revision of the family Carabodidae (Acari: Oribatida) III. Redefinition of Meriocepheus peregrinus Aoki, 1973; Bathocepheus concavus Aoki, 1978; and Opisthocepheus kirai Aoki, 1976. International Journal of Acarology 39(4): 327-340. doi:10.1080/01647954.2013.781675

38. Fernandez N., Theron P.D., Rollard C. 2013. Revision of the family Carabodidae (Acari: Oribatida) IV. Afticarabodes anjavidilavai gen.nov., sp.nov., Rugocepheus joffrevillei sp.nov. and redefinition of the genus Rugocepheus Mahunka, 2009. International Journal of Acarology 39 (6): 462-480. doi:10.1080/01647954.2013.822928

39. Fernandez N., Theron P.D., Rollard C., Leiva S. 2013. The family Carabodidae (Acari: Oribatida) V. The genus Congocepheus (first part) with redescriptions of Congocepheus heterotrichus Balogh, 1958, Congocepheus orientalis Mahunka, 1987 and Congocepheus hauseri Mahu8nka, 1989. International journal of Acarology 39(8): 600-614. doi:10.1080/01647954.2013.858771

40. Fernandez N., Theron P.D., Rollard C., Tiedt L.R. 2013. The family Carabodidae (Acari: Oribatida) V. The genus Congocepheus Balogh, 1958 (second part), with a redescription of Congocepheus involutus Mahunka, 1997, and descriptions of two new species. Zoosystema 35(4): 551-579. doi:10.5252/z2013n4a8

41. Leiva S., Fernandez N.,Theron P., Rollard C. 2013. Agistemus aimogastaensis sp.n. (Acari, Actinedida, Stigmaeidae), a recently discovered predator of eriophyid mites Aceria oleae and Oxycenus maxwelli, in olive orchards in Argentina. ZOOKEYS 312: 65-78. doi:10.3897/zookeys.312.5520

42. Fernandez N.,Theron P., Rollard C. 2013. First discovery of Plasmobatidae (Acari, Oribatida) in Gabon, redefinition and new species of the genus Solenozetes Grandjean, 1932. Zoosystema 35(2): 137- 150. doi:10.5252/z2013n2a1

43. Jubileus M.T., Theron P.D., Van Rensburg L., Maboeta M.S. 2013. Utilizing Eisenia Andrei to assess the ecotoxicity of platinum mine tailings disposal facilities. Ecotoxicology 22: 331-338. doi:10.1007/s10646-012-1029-6

44. Fernandez N., Theron P.D., C. Rollard C., Castillo E.R. 2014. Revision of the family Carabodidae (Acari, Oribatida) V (third part). Redefinition of Congocepheus, definition of Cavaecarabodes gen. nov., and descriptions of three new species, Congocepheus germani sp.nov., Cavaecarabodes pulchritude gen.nov., sp. nov., and Cavaecarabodes anouchkae gen. nov., sp. nov. International Journal of Acarology 40(7): 535-555. doi:10.1080/01647954.2014.959050

45. Fernandez N., Theron P.D., Leiva S., Rollard C., Tiedt L.R. 2014. Revision of the family Carabodidae (Acari: Oribatida) VI. Mangabebodes kymatismosi gen.nov, sp.nov. and Antongilibodes paulae gen.nov., sp.nov. from Madagascar. International Journal of Acarology 40(4): 296-319. doi:10.1080/01647954.2014.914972

46. Fernandez N., Theron P.D., Rollard C., Castillo E.R. 2014. Revision of the family Carabodidae (Acari, Oribatida) VII. Redefinition of the genus Malgasodes; redescription of M. curvisetus Mahunka and complementary description of M. hungarorum Mahunka. Phylogenetic relationships between Malgasodes, Bovicarabodes, Afticarabodes, Congocepheus and Cavaecarabodes are discussed. ZooKeys 435: 25-48. doi:10.3897/zookeys.435.8071

47. Fernandez N., Theron P.D., Rollard C., Leiva S. 2014. The family Carabodidae (Acari: Oribatida) VIII. The genus Machadocepheus (first part) Machadocepheus leoneae sp. nov. and Machadocepheus rachii sp. nov. from Gabon. ZooKeys 456: 1-28. doi:10.3897/zookeys.456.8570

48. Fernandez N., Theron P.D., Rollard C., Castillo E.R. 2014. Oribatid mites from deep soils of HònChông limestone hills of Kien Giang Province, Vietnam. I. The family Lohmanniidae (Acari: Oribatida), first part. Bedoslohmannia anneae n. gen., n. sp., and Paulianacarus vietnamese n. sp. Zoosystema 36(4): 771- 787. doi:10.5252/z2014n4a5

49. Fernandez N., Coineau Y., Theron P.D., Tiedt L.R. 2014. Nannodromus reveilleti (Acari, Anystida, Saxidromidae) a new genus from South Africa. ZooKeys 378: 17-39. doi:10.3897/zookeys.378.6753

50. Fernandez N., Theron P.D., Leiva S. 2016. Revision of the family Carabodidae (Acari, Oribatida) V. Fourth part. Two new species of the genus Congocepheus from the Republic of Rwanda: Congocepheus rwandensis sp. n., and Congocepheus kayoveae sp. n. ZooKeys 556: 19-41. doi:10.3897/zookeys.556.7011

51. Fernandez N., Theron P.D., Rollard C., Leiva S. 2015. Oribatid mites (Acari: Oribatida) from deep soils of Hὁn Chông limestone hills, Kien Giang Province, Vietnam. II. Descriptions of two new species, Papillacarus whitteni sp. nov. (family Lohmannidae) and Basilobelba maidililae sp. nov. (family Basilobelbidae). International Journal of Acarology, 41(2): 132-146. doi:10.1080/01647954.2015.1014414

52. Fernandez N., Theron P.D., Rollard C., Leiva S. 2015. The family Carabodidae (Acari: Oribatida) IX. Redescription of Carabodella calcarata Mahunka, 1986 and Hardybodes penicillatus Mahunka, 1995. International Journal of Acarology 41(7): 551-562. doi:10.1080/01647954.2015.1081406

53. Fernandez N., Theron P.D., Rollard C., Leiva S., Tiedt L.R. 2016. Revision of the family Carabodidae (Acari: Oribatida) XI. Congocepheus kardiae sp. nov. and Zimbabweae pluosiae gen. nov., sp. nov. from the Republic of Zimbabwe. International Journal of Acarology 42(7): 341-357. doi:10.1080/01647954.2016.1199138

54. Fernandez N., Theron P. D., Rollard C., Leiva S. 2016. Revision of the family Carabodidae (Acari: Oribatida) X. Bovicarabodes jacquelinae sp.nov., redescription of the genus Tuberocepheus Balogh & Manhunka, 1969 and redescription of Tuberocepheus longus (Balogh, 1962). International Journal of Acarology 42(2): 79-91. doi:10.1080/01647954.2015.1124920

55. Janion-Scheepers C., Measey J., Braschler B., Chown S.L., Coetzee L., Colville J.F., Dames J., Davies A.B., Davies S.J., Davis A.L.V., Dippenaar-Schoeman A.S., Duffy G.A., Fourie D., Griffiths C., Haddad C.R., Hamer M., Herbert D.G., Hugo-Coetzee E.A., Jacobs A., Jacobs K., Jansen van Rensburg C., Lamani S., Lotz L.N., Louw S. vdM., Lyle R., Malan A.P., Marais M., Neethling J.-A., Nxele T.C., Plisko D.J., Prendini L., Rink A.N., Swart A., Theron P.D., Truter M., Ueckermann E.A., Uys V., Villet M. H., Willows-Munro S., Wilson J.R.U. 2016. Soil biota in a megadiversity country: Current knowledge and future research directions in South Africa. Pedobiologia-Journal of Soil Ecology 59: 129-174. doi:10.1016/j.pedobi.2016.03.004

56. Fernandez N., Theron P.D., Leiva S., Tiedt L.R. 2017. Two new oribatid mites from Costa Rica Mixacarus turialbaiensis sp.n. and Paulianacarus costaricensis sp.n. (Acari, Oribatei, Lohmannidae). ZooKeys 680: 33-56. doi:10.3897/zookeys.680.13213

57. Ermilov S.G., Hugo-Coetzee E.A., Theron P.D. 2017. New and interesting moss mites (Acari: Oribatida) from South Africa, with description of two new species. Systematic and Applied Acarology 22(10): 1560-1573. doi:10.11158/saa.22.10.2

58. Ermilov S.G., Hugo-Coetzee E.A. Khaustov A.A. & Theron P.D. 2017. New and interesting mites (Acari: Oribatida) near Potchefstroom (South Africa), with description of two new species. Systematic and Applied Acarology 22(11): 1849-1871. doi:10.11158/saa.22.11.6

59. Ermilov S.G., Hugo-Coetzee E.A., Theron P.D., Behan-Pelletier V.M. 2017. Contribution to the knowledge of the oribatid mite family Nesozetidae (Acari: Oribatida). Zootaxa 4358: 311-327. doi:10.11646/zootaxa.4358.2.5

60. Ermilov S.G., Hugo-Coetzee E.A. Khaustov A.A., Theron P.D. 2017. Juvenile instars of alumna curvifamulus (Acari, Oribatida, Galumnidae). Acarina 25(2): 87-95. doi:10.21684/0132-8077-2017-25-2-87-95

61. Khaustov A.A., Ueckermann E.A., Theron P.D. 2017. A new species of Stigmaeus (Acari: Prostigmata: Stigmaeidae) from South Africa. Systematic and Applied Acarology 22(9): 1413-1421. doi:10.11158/saa.22.9.8

62. Fernandez N., Theron P.D., Leiva S., Tiedt L.R. 2017. Revision of the family Carabodidae (Acari, Oribatida) XII. Yoshiobodes camerunensis sp.n. and Rugocepheus costaricensis sp.n. ZooKeys 706: 31-50. doi:10.3897/zookeys.706.14807

63. Fernandez N., Theron P.D., Leiva S., Tiedt L.R. 2017. Revision of the family Carabodidae (Acari, Oribatida) XIII. Congocepheus camerunensis sp.n. from Cameroon and Synkrotima zimbabwea gen.nov., sp.nov. from the Republic of Zimbabwe. Systematic and Applied Acarology 22(11): 1822-1842. doi:10.11158/saa.22.11.4

64. Fernandez N., Theron P.D., Leiva S., Tiedt L.R. 2017. Revision of the family Carabodidae (Acari, Oribatida) XIV. Phyllocarabodes costaricensis sp. nov. from Costa Rica and Zimbabweae kenyaensis sp. nov. from Kenya. International Journal of Acarology 43(7): 1-16. doi:10.1080/01647954.2017.1366549

65. Fernandez N., Theron P.D., Leiva S., Jordaan A. 2018. Revision of the family Carabodidae (Acari, Oribatida) XV. Costacarabodes turrialbai gen. nov., and Tuberocepheus kompsosis sp. nov. from Costa Rica. International Journal of Acarology 44(6): 236-253. doi:10.1080/01647954.2018.1508249

66. Fernandez N., Theron P.D., Leiva S., Jordaan A. 2018. The family Lohmammiidae (Acari, Oribatida) II: two new Oribatid mites, Meristacarus perikopesis sp. nov. from Costa Rica and Torpacarus eidikoterai sp. nov. from Kenya. Zookeys (743): 43-66. doi:10.3897/zookeys.743.22815

67. Ermilov S.G., Hugo-Coetzee E.A., Theron P.D. 2018. To the knowledge of Oribatid mites of the subgenus Galumna (Galumna) Heyden (Acari, Oribatida, Galumnidae) in South Africa, with a key to species known from the Ethiopian Region. Zoologicheskii Zhurnal 97(5): 515-527. doi:10.7868/S0044513418050033

68. Ueckermann E.A., Palevsky E., Gerson U., Recht E., Theron P.D. 2018. The Tenuipalpidae (Acari: Trombidiformes) of Israel. Acarologia 58: 483-525.

69. Khaustov A.A., Hugo-Coetzee E.A., Ermilov S.G., Theron P.D. 2019. A new genus and species of the family Microdispidae (Acari: Heterostigmata) associated with Trinervitermes trinervoides (Sjöstedt) (Isoptera: Termitidae) from South Africa. Zootaxa 4647(1): 104-114. doi:10.11646/zootaxa.4647.1.10

70. Ermilov S.G., Hugo-Coetzee E.A. Khaustov A.A., Theron P.D. 2019. Oribatid mites (Acari: Oribatida) inhabiting termite nests in the Faan Meintjes Nature Reserve (South Africa). Systematic and Applied Acarology 24(9): 1783-1798. doi:10.11158/saa.24.9.14

71. McGuirk B., Theron P.D., Maboeta M. 2020. The effects of different gold mine tailings on growth, reproduction and avoidance-behaviour of earthworms. African Zoology 55: 35-42. doi:10.1080/15627020.2020.1724826

72. Uusitalo M., Ueckermann E.A., Theron P.D. 2020. A review of the family Alycidae (Acari, Acariformes) from South Africa. Zootaxa 4858 (3): 301-340. doi:10.11646/zootaxa.4858.3.1

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